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Rieppel, Olivier, Zaher, Hussam, Tchernov, Eitan, Polcyn, Michael J. (2003) The anatomy and relationships of Haasiophis terrasanctus, a fossil snake by well-developed hind limbs from the Mid-Cretaceous of the Middle East. Journal of Paleontology, 77 (3) 536-558 doi:10.1017/s0022336000044231

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Reference TypeJournal (article/letter/editorial)
TitleThe anatomy and relationships of Haasiophis terrasanctus, a fossil snake by well-developed hind limbs from the Mid-Cretaceous of the Middle East
JournalJournal of Paleontology
AuthorsRieppel, OlivierAuthor
Zaher, HussamAuthor
Tchernov, EitanAuthor
Polcyn, Michael J.Author
Year2003 (May)Volume77
Issue3
PublisherCambridge University Press (CUP)
DOIdoi:10.1017/s0022336000044231Search in ResearchGate
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Mindat Ref. ID419745Long-form Identifiermindat:1:5:419745:8
GUID0
Full ReferenceRieppel, Olivier, Zaher, Hussam, Tchernov, Eitan, Polcyn, Michael J. (2003) The anatomy and relationships of Haasiophis terrasanctus, a fossil snake by well-developed hind limbs from the Mid-Cretaceous of the Middle East. Journal of Paleontology, 77 (3) 536-558 doi:10.1017/s0022336000044231
Plain TextRieppel, Olivier, Zaher, Hussam, Tchernov, Eitan, Polcyn, Michael J. (2003) The anatomy and relationships of Haasiophis terrasanctus, a fossil snake by well-developed hind limbs from the Mid-Cretaceous of the Middle East. Journal of Paleontology, 77 (3) 536-558 doi:10.1017/s0022336000044231
In(2003, May) Journal of Paleontology Vol. 77 (3) Cambridge University Press (CUP)
Abstract/NotesThe fossil snake species Haasiophis terrasanctus Tchernov, Rieppel, Zaher, Polcyn, and Jacobs, 2000, from the early Upper Cretaceous of the Middle East, is described and illustrated, following a review of the current debate on snake relationships and origins. The description and discussion presented here adds important detail to the knowledge of this taxon and its phylogenetic significance beyond the limited account presented in the original description of Haasiophis. The species is remarkable in that it shows the skull of a relatively advanced (i.e., macrostomatan) snake, yet preserves well-developed hind limbs. The hind limb includes a femur, tibia, fibula, astragalus, calcaneum, distal tarsal four, and remains of four metatarsals and two phalanges. Haasiophis cannot be considered a juvenile specimen of Pachyrhachis. The implications of the presence of well-developed hind limbs in Haasiophis, Pachyrhachis, and Podophis for the cladistic analysis of the phylogenetic interrelationships of these fossil snakes is discussed. The presence of well-developed hind limbs in Pachyrhachis and Haasiophis also creates methodological problems for the cladistic analysis of the phylogenetic relationships of these fossil snakes. Scenarios of snake origins are reviewed and found to be deficient in the absence of a well-corroborated hypothesis of snake relationships within Squamata.


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